From Wikipedia, the free encyclopedia

The leopard; (Panthera pardus) is an Old World mammal of the Felidae family and the smallest of the four big cats in the genus Panthera, the other three are the tiger, lion and jaguar. Once distributed across southern Asia and Africa, from Korea to South Africa, the leopard's range of distribution has decreased radically over time because of a variety of factors, including human influence, and the leopard now chiefly occurs in sub-Saharan Africa. There are fragmented populations in India, Indochina, Malaysia, and China. Despite the loss of range and continual declines in population, the cat remains a "Least Concern" species; its numbers are greater than that of the other Panthera species, all of which face more acute conservation concerns.

The leopard has relatively short legs and a long body, with a large skull. Physically, it most closely resembles the jaguar, although it is usually smaller and of slighter build. Its fur is marked with rosettes which lack internal spots, unlike those of the jaguar. Leopards that are melanistic, either completely black or very dark in coloration, are one of the big cats known colloquially as black panthers.

The species' success in the wild owes in part to its opportunistic hunting behaviour, its adaptability to a variety of habitats and its ability to move at up to approximately 60 kilometres (37 miles) an hour. The leopard consumes virtually any animal it can hunt down and catch. Its preferred habitat ranges from rainforest to desert terrains. Its ecological role and status resembles that of the similarly-sized cougar in the Americas.

The leopard is an agile and stealthy predator. Although smaller than the other members of the Panthera genus, the leopard is still able to take large prey given a massive skull that well utilizes powerful jaw muscles. Its body is comparatively long for a cat and its legs are short. Head and body length is between 90 and 190 cm (35 and 75 in), the tail reaches 60 to 110 cm (24 to 43 in). Shoulder height is 45 to 80 cm (18-31 in). Males are considerably larger than females and weigh 37 to 91 kg (82 to 200 lbs) compared to 28 to 60 kg (62 to 132 lbs) for females. The larger-bodied populations of leopard (such as the Javan leopard and the leopards from the forested mountains and tropical rainforests of Africa) are generally found in areas isolated from competing large predators, especially from dominant big cats like Lions and Tigers.

One of many spotted cats, a leopard may be mistaken for a cheetah or a jaguar (though, of course, there is no range overlap with the latter). The leopard has rosettes rather than cheetah's simple spots, but they lack internal spots, unlike the jaguar. The leopard is larger and less lanky than the cheetah but smaller than the jaguar. The leopard's black, irregular rosettes serve as camouflage. They are circular in East Africa but tend to be square-shaped in southern Africa.

Leopards have been reported to reach 21 years of age in captivity.

Biology and behavior

Diet and hunting

Leopards are opportunistic hunters. Although mid-sized animals are preferred, the leopard will eat anything from dung beetles to 900 kg (1,984 lb) male giant elands. Their diet consists mostly of ungulates and monkeys, but rodents, reptiles, amphibians, birds and fish are also eaten. In Africa, mid-sized antelopes provide a majority of the leopard's prey, especially impala and Thomson's gazelles. In Asia the leopard preys on deer such as chitals and muntjacs as well as various Asian antelopes and Ibex. One study at the Wolong Reserve in China revealed how adaptable the leopard's hunting behaviour is: over the course of seven years vegetative cover receded, and the animals opportunistically shifted from primarily consuming tufted deer to instead pursuing bamboo rats and other smaller prey.

The leopard stalks its prey silently and at the last minute pounces on its prey and strangles its throat with a quick bite. Leopards often hide their kills in dense vegetation or take them up trees, and are capable of carrying animals up to three times their own weight this way.

One survey of nearly 30 research papers found preferred prey weights of 10 to 40 kg (22-88 lb), with 25 kg (55 lb) most preferred. Along with impala and chital, a preference for bushbuck and common duiker was found. Other prey selection factors include a preference for prey in small herds, in dense habitat, and those that afford the predator a low risk of injury.

Social structure and home range

Studies of leopard home range size have tended to focus on protected areas, which may have led to skewed data; as of the mid-1980s, only 13% of the leopard range actually fell within a protected area. In their IUCN survey of the literature, Nowell and Jackson suggest male home territories vary between 30-78 square kilometers, but just 15-16 km² for females. Research in a conservation area in Kenya shows similar territory sizes and sex differential: 32.8 km² ranges for males, on average, and 14 km² for females. In Nepal, somewhat larger male ranges have been found at about 48 km², while female ranges are in-keeping with other research, at 17 km²; female home territories were seen to decrease to just five to seven km² when young cubs were present, while the sexual difference in range size seemed to be in positive proportion to overall increase. However, significant variations in size of home territories have been suggested across the leopard's range. In Namibia, for instance, research that focussed on spatial ecology in farmlands outside of protected areas found ranges that were consistently above 100 km², with some more than 300 km²; admitting that their data were at odds with others', the researchers also suggested little or no sexual variation in the size of territories. Virtually all sources suggest that males do have larger ranges. There seems to be little or no overlap in territory amongst males, although overlap exists between the sexes; one radio-collar analysis in the Ivory Coast found a female home range completely enclosed within a male's.

The leopard is solitary and, aside from mating, interactions between individuals appear to be infrequent. Aggressive encounters have been observed, however. Two of five males studied over a period of a year at a game reserve in South Africa died, both violently. One was initially wounded in a male-male territorial battle over a carcass; taken in by researchers, it was released after a successful convalescence only to be killed by a different male a few months later. A second was killed by another predator, possibly a spotted hyena. A third of the five was badly wounded in intraspecific fighting, but recovered.

Ecology

Distribution and habitat

Data from 1996 found that the leopard has the largest distribution of any wild cat, occurring most in certain parts of southern Asia and widely in eastern and central Africa, although populations before and since have shown a declining trend and are fragmented outside of subsaharan Africa. The IUCN notes that within sub-Saharan Africa the species is "still numerous and even thriving in marginal habitats" where other large cats have disappeared, but that populations in North Africa may be extinct. In Asia, data on distribution is not consistent: populations in Southwest and Central Asia are small and fragmented; in the northeast portion of the range, they are critically endangered; but in the Indian subcontinent, Southeast Asia, and China, the leopard is still relatively abundant.

Leopards live mainly in grasslands, woodlands and riverside forests. The animal has primarily been studied in open savannah habitats, which may have biased common descriptions. It is generally considered nocturnal, for instance, but radio-tracking and scat analysis in West Africa has found that rainforest leopards are more likely to be diurnal and crepuscular. Forest leopards are also more specialized in prey selection and exhibit seasonal differences in activity patterns. While associated with the savanna and rainforest, the leopard is exceptionally adaptable: in the Russian Far East, the animal inhabits temperate forests where winter temperatures reach a low of –25 °C.

Ecological role

Because of their wide habitat range, leopards must compete for food and safety with other large predators such as lions, tigers, spotted hyenas and wild dogs. These competitors sometimes may steal the leopard's kill or devour its young. A single lion or tiger is capable of killing an adult leopard. Leopards have adapted to live alongside these other predators by hunting at different times of the day, and by avoiding areas frequented by them. In search of safety, the leopard will often stash its young or a recent kill high up in a tree. Lions are occasionally successful in climbing trees and fetching leopard kills, and if motivated, an adult tiger might also scale a tree to acquire food.

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